Research

Peer-reviewed Research

Lobón-Rovira, J., Vaz Pinto, P., Keates, C., & Stanley, E. L. (2025). Ancient polymorphism, secondary contact, taxonomic inflation or all of them together? Mito-nuclear discordance in the Angolan girdled lizard (Cordylidae: Cordylus), with the description of a new species from the central highlands. Zoological Journal of the Linnean Society, 205(1), zlaf112.

Mungenge, C. P., Wasserman, R. J., Mlambo, M. C., Dondofema, F., Keates, C., Bute, T. F., & Dalu, T. (2025). Diversity of large branchiopods (Anostraca, Notostraca, Spinicuadata) in semi-arid temporary pans of North West Province, South Africa, including a new record for South Africa. Journal of Crustacean Biology, 45(2), ruaf032.
Conradie, W., Hundermark, C., Kemp, L., & Keates, C. (2025). New Pygmy Gecko (Goggia: Gekkonidae) from the arid Northern Cape Province of South Africa. Zootaxa, 5618(4), 552-570.
Leshaba, E. N., Dube, T., Dondofema, F., Munyai, L. F., Keates, C., Riddell, E., … & Dalu, T. (2025). Macroinvertebrate diversity within pan wetlands in relation to geological type and hydroperiod in a protected subtropical Austral national park. Chemistry and Ecology, 41(5), 587-609.
Keates, C., Wasserman, R. J., Conradie, W., Dondofema, F., Munyai, L., Riddell, E., & Dalu, T. (2024). Frogs of the Makuleke Contractual Park, northern Kruger National Park. Koedoe, 66(1), 9.
Keates, C., Conradie, W., & Edwards, S. (2024). Phylogeographic structuring in a widespread southern African grass snake (Psammophiinae: Psammophylax rhombeatus). African Journal of Herpetology, 73(2), 210-235.
Cowan, O. S., Conradie, W., Keates, C., Telford, N. S., Petford, M. A., Egan, V., & Tolley, K. A. (2025). Lost lizards: The importance of targeted surveys in filling knowledge gaps for reptile conservation in South Africa. Journal for Nature Conservation, 84, 126790.
Dalu, T., Chari, L. D., Keates, C., Munyai, L. F., Mutoti, M. I., & Mpopetsi, P. P. (2025). Rivers of North Africa. In Afrotropical Streams and Rivers (pp. 163-193). Elsevier.
Dalu, T., Masese, F. O., Parker, D. M., Chanyandura, A., Dube, T., Fugère, V., … & Mpopetsi, P. P. (2025). Overview and future prospects of African rivers research. In Afrotropical Streams and Rivers (pp. 773-785). Elsevier.
Keates, C., & Reid, M. K. (2025). Birds, mammals, reptiles, and amphibians. In Afrotropical Streams and Rivers (pp. 497-518). Elsevier.
Bute, T. F., Wyness, A., Wasserman, R. J., Dondofema, F., Keates, C., & Dalu, T. (2024). Microbial community and extracellular polymeric substance dynamics in arid–zone temporary pan ecosystems. Science of The Total Environment, 932, 173059.
Chaney, T., Pauwels, O. S., Nagy, Z. T., Gvoždík, V., Kusamba, C., Badjedjea, G., … & Greenbaum, E. (2024). Phylogenetics and Integrative Taxonomy of African Water Snakes (Squamata: Colubridae: Grayia). Herpetological Monographs, 38(1), 1-52.
Munyai, L. F., Mugwedi, L., Wasserman, R. J., Dondofema, F., Riddell, E., Keates, C., & Dalu, T. (2024). Water and sediment chemistry drivers of chlorophyll-a dynamics within a Ramsar declared floodplain pan wetland system. Environmental Science and Pollution Research, 31(19), 28549-28563.
Wasserman, R. J., Dondofema, F., Keates, C., Cuthbert, R. N., & Dalu, T. (2023). Turtle-mediated dispersal of anostracan dormant eggs: Evidence for dominance hierarchy effects. Ecology, 104(7), e4066.
Baptista, N. L., Pinto, P. V., Keates, C., Lobón-Rovira, J., Edwards, S., & Rödel, M. O. (2023). Two new Poyntonophrynus species (Anura: Bufonidae) highlight the importance of Angolan centers of endemism. Vertebrate Zoology, 73, 991-1031.
Munyai, L. F., Mugwedi, L., Wasserman, R. J., Dondofema, F., Riddell, E., Keates, C., & Dalu, T. (2023). Metal and non–metal dynamics and distribution in soil profiles across selected pans in the Ramsar declared subtropical within a national protected area. Chemistry and Ecology, 1-21.
Mungenge, C. P., Wasserman, R. J., Dondofema, F., Keates, C., Masina, F. M., & Dalu, T. (2023). Assessing chlorophyll–a and water quality dynamics in arid–zone temporary pan systems along a disturbance gradient. Science of The Total Environment, 873, 162272.
Masina, F. M., Wasserman, R. J., Wu, N., Mungenge, C. P., Dondofema, F., Keates, C., … & Dalu, T. (2023). Macroinvertebrate diversity in relation to limnochemistry in an Austral semi–arid transboundary aquifer region pan system. Science of The Total Environment, 878, 163161.
Conradie, W., Keates, C., Verburgt, L., Baptista, N. L., & Harvey, J. (2023). Contributions to the herpetofauna of the Angolan Okavango-Cuando-Zambezi river drainages. Part 3: Amphibians.
Keates, C., Conradie, W., Dalu, T., Dondofema, F., Riddell, E. S., & Wasserman, R. J. (2022). Phylogenetic placement of the enigmatic Floodplain water snake, Lycodonomorphus obscuriventris FitzSimons, 1964. Koedoe, 64(1), 1-9.

Cuthbert RN, Wasserman RJ, Keates C and Dalu T. (2022). Food webs. Dalu T & Wasserman RJ (Eds.). Fundamentals of Tropical Freshwater Wetlands: From Ecology to Conservation Management (pp. 517-547). Elsevier.
Conradie, W., Keates, C., Verburgt, L., Baptista, N. L., Harvey, J., Julio, T., & Neef, G. (2022). Contributions to the herpetofauna of the Angolan Okavango-Cuando-Zambezi River drainages. Part 2: Lizards (Sauria), chelonians, and crocodiles.
Baptista N.L, Pinto PV, Keates C., Edwards S., Roedel MO., & Conradie W. (2021). A new species of red toad, Schismaderma Smith, 1849 (Anura: Bufonidae), from central Angola. Zootaxa 5081(3), 301-332.
Lobon-Rovira J, Conradie W, Vaz Pinto P, Keates C, Edwards S, Du Plessis A, & Branch WR. (2022). Systematic revision of Afrogecko ansorgii (Boulenger, 1907) (Sauria: Gekkoni-dae) from western Angola. Zootaxa.
Conradie, W., Keates, C., Baptista, N. L., & Lobón-Rovira, J. (2022). Taxonomical review of Prosymna angolensis Boulenger, 1915 (Elapoidea, Prosymnidae) with the description of two new species. ZooKeys 1121: 97–143.
Conradie, W., Keates, C., Lobón-Rovira, J., Pinto, P. V., Verburgt, L., Baptista, N. L., … & Júlio, T. (2020). New insights into the taxonomic status, distribution and natural history of De Witte’s Clicking Frog (Kassinula wittei Laurent, 1940). African Zoology, 55(4), 311-322.
Conradie, W., Baptista, N. L., Verburgt, L., Keates, C., Harvey. J., Júlio, T., Neef, G. (2021). Contributions to the herpetofauna of the Angolan Okavango: Cuando-Zambezi River drainages. Part 1: Serpentes (snakes). Amphibian & Reptile Conservation 15(2) [General Section]: 244–278 (e292).
Nicolau, G. K., Petford, M., Edwards, S., Busschau, T., Lynch, K., Kemp, L., Balmer, J., Keates, C., Hundermark, C. R., Weeber, J., Conradie, W. (2021). New insights into the geographical distribution, ecology and conservation status of South Africa’s endemic Coastal Leaf-toed Gecko, Cryptactites peringueyi (Boulenger, 1910), Herpetology Notes, 14, 439-450.
Branch, W. R., Baptista, N., Keates, C., & Edwards, S. (2019). Rediscovery, taxonomic status, and phylogenetic relationships of two rare and endemic snakes (Serpentes: Psammophiinae) from the southwestern Angolan plateau. Zootaxa, 4590(3), 342-366.
Keates, C., Conradie, W., Greenbaum, E., & Edwards, S. (2019). A snake in the grass: Genetic structuring of the widespread African grass snake (Psammophylax Fitzinger 1843), with the description of a new genus and a new species. Journal of Zoological Systematics and Evolutionary Research, 57(4), 1039-1066.
Conradie, W., Deepak, V., Keates, C., & Gower, D. J. (2020). Kissing cousins: a review of the African genus Limnophis Günther, 1865 (Colubridae: Natricinae), with the description of a new species from north-eastern Angola. African Journal of Herpetology, 69(1), 79-107.

Popular Science Book Chapters

Branch, B., & Keates, C. (2021). Reptiles and Amphibians around Grahamstown. In Guide to the Natural and Cultural History of Grahamstown / Makhanda, edited by Lubke, R., & De Moor, I.

Conferences

13th Herpetological Association of Africa Conference (2017). Hluhuwe, RSA. Poster presentation

Joint SANBI & FBIP Forum (2018). St Francis, RSA. Oral presentation

14th Herpetological Association of Africa Conference (2019). St Francis, RSA. Oral Presentation and mini oral presentation

9th World Congress of Herpetology Conference (2020). Dunedin, NZ. Oral Presentation

Southern African Society of Aquatic Scientists Conference (2021). Virtual conference. Oral presentation

Herpetological Association of African Mini Conference (2021). Virtual Conference. Oral Presentation

Southern African Society of Aquatic Scientists Conference (2022). Bloemfontein, RSA. Oral presentation

15th Herpetological Association of Africa Conference (2023). Hoedspruit, RSA. Two Oral Presentations

Close–out Workshop for the Groundwater Dependent Ecosystems (GDEs) and Biodiversity in Khakea/Bray Transboundary Aquifer Project (2023). Mafikeng, RSA. Oral Presentation (Virtually presented).

Florida Aquatic Plant Society Conference (2024). Daytona, USA. Oral Presentation

Florida Chapter of the Wildlife Society Spring Conference (2025). Tallahassee, USA. Oral Presentation

UF Aquatic Weed Control Short Course (2025) . Orlando, USA, Two Oral Presentations

17th International Symposium on Aquatic Plants (2025). Lisbon, Portugal. Oral Presentation

Florida Aquatic Plant Society Conference (2025). St Petersburg, USA. Oral Presentation


Services Provided

Scientific Consulting

Reptile and Amphibian Guide

Reptile and Amphibian Presentations, Talks and Walks

Stock Photography

Snake Removal

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Chad Keates

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